Regarding anatomy, yep, I still have work I'm starting from nothing (those are the first humans I ever drew seriously) and I'm learning as I draw, not much time to do anatomy studies unfortunately... I should try to find time...
The comment on pacing is super interesting to me. I was a bit aware there MAY be an issue but no one ever said anything so I wasn't sure if I was imagining it or not. That's exactly why these threads are useful whether some like it or not! It is a voluntary thing to give a non-linear and somewhat random structure to my comic, but it should not be difficult to follow, so this is a very good info to get, that it is indeed difficult to follow. I think it gets worse in later chapters too, so certainly something I have to work on, Thanks again.
Now Runner. The things I liked less are rather at the beginning, so I suppose it's less relevant but I'll still tell you.
First, I feel you need a page or at least a panel before the very first panel. Maybe an environment/exposition panel. I was surprised it was starting so abruptly with a relatively small panel with text, so I scrolled up trying to find the first page before realizing that was it. It is understandable, but a bit abrupt.
Also, I tend to really not like action panels with completely empty backgrounds. I know it is to keep the focus on the character but I feel it looks unfinished. Just a shadow or a blurry simple environmental object would be enough. But, I saw you started changing that later on so maybe not an issue at all.
Last things that seems to remain, but I guess it is an artistic choice so... the 3/4 faces with the face drawn entirely like if it was front facing (or if it was almost front facing) are a bit disturbing to me. It's super common in comics - especially manga- so maybe not an issue for most but as the topic is what I don't like.. I don't like that
I very much enjoyed the aggressive fauna. I was super surprised by the land bobbit because I have a land bobbit too. The ultimate animal horror as far as I'm concerned