Hi there! I'll do my best to help. Constructive critique incoming!
I think your biggest strength is composition. The action is generally easy to follow, but you have a nice mix of longer shots and close-ups, and you play around with some interesting camera angles. Composition is way more important for a visual storytelling medium than making each individual panel look pretty, so kudos. Your characters are also visually distinct (making them animals helps, of course) and their proportions stay relatively consistent, which is great.
As for stuff to improve, I'd say the biggest thing you can do is rely more on reference. I like how you don't shy away from drawing backgrounds, but they don't seem to have the right textures. Mud and rocks are drawn with squiggly lines that don't really communicate what mud or rock feel or look like. Trees look strangely squishy, as if they were topped with dollops of whipped cream, not masses of leaves. Splashing water on pg. 12 was given the same spiky lines as the flames a page earlier. An explosion is drawn with a texture that looks similar to mud; if you hadn't coloured it orange I wouldn't have known what it was! I would recommend looking up photos of what you want to draw before you draw it -- not to copy, but to get a stronger idea of what something really looks like, not what you think it looks like. Draw practice studies when you have time.
I'd also focus on the linework. It's a little weak, a little thin, and a little unsure. Don't be afraid to add some bolder, more energetic strokes, and vary the line width a little. A character's outline should be drawn a little thicker than smaller details, like clothing folds or scales. Lines in shadow should be drawn a bit thicker than lines facing the light, and lines for foreground details should be much thicker than lines for background details. The weak, unsure linework may partially explain why your poses are a little stiff. Gesture studies may help inject some energy into your art -- https://vimeo.com/croquiscafe has a free collection of videos of nude models for artists; I've been relying on it a lot to study both gesture and human anatomy. Don't forget to study animal anatomy too, of course
Your shading has improved the most with time, which has given your newer pages much more depth. However, you still rely a little too much on soft gradients and soft, smudgy shadows. Some soft lighting is fine, if the light source is weak and far away, but don't be afraid to use bolder shadows with sharper edges. Bolder, sharper lights and shadows will really make your characters pop.
In summary: use more reference, and have more confidence, both in your lines and in your lighting. I hope my novel helped at least a little. All in all, I can see that you have a lot of passion for your story, and it's shaping up to be a grand one!